A thought:

It was only when the early multi-cellular aggregators developed characteristics which, when realized and expressed through the aggregate, outweighed in benefit to the genome the harm to the genome those characteristics represented by the loss of efficient independent action, that the morality of the multi-cellular aggregators began to call for perfection.

It was only when the first perfected multi-cellular moralities were realized that multi-cellular life was fully realized.

One aspect of a perfected multi-cellular morality is that no part lives for itself, instead every part lives for the whole, all of its action directed towards the common good. The part cannot conceive of the whole and decide how to be, it must simply follow the rule of its perfected morality, crystalized in its structure, one manifestation of the greater morality, the greater genome.

In an unperfected multi-cellular morality, the parts sometimes act for themselves to the detriment of all, they do not conceive of the whole and resist it, they simply follow their rule and manifest their imperfect morality.

What makes their morality imperfect is the very fact itself of the characteristics that have developed such that the benefit those characteristics bring to the genome when expressed as part of the aggregate outweigh the harm to the genome represented in the loss of efficiency in independent action those characteristics bring the individual part.

Can we see where this goes?

A hope:

The human techno-cultural capacity is an evolutionary ratcheting mechanism.